best chemical biology phd programs

Chemical Biology, PhD

Zanvyl krieger school of arts and sciences, admission requirements.

Students interested in pursuing multidisciplinary research to gain a broad perspective in chemistry and biology are encouraged to apply to the Chemistry-Biology Interface (CBI) PhD program. The CBI Program spans departments in Johns Hopkins University’s schools of Arts and Sciences, Engineering, Medicine, and Public Health.

Selected applicants will be asked to visit Johns Hopkins University for an interview the first week of March 2024. The CBI Program will arrange for meetings with faculty and students, and conduct tours of the facilities and community.

Application Instructions

To apply to the CBI Program, you will need to submit:

-  An online application  for the Chemistry-Biology Interface (CBI) PhD - An unofficial undergraduate transcript - Statement of purpose - Three (3) letters of recommendation

In addition, the general GRE along with one of the following subject tests are  optional  but may be submitted:

- Biochemistry - Cell and Molecular Biology - Biology - Chemistry

International Applicants

A training grant from the National Institutes of Health provides support for our students during their first year in the graduate program at the Chemistry-Biology Interface. However, the NIH limits this to U.S. citizens and permanent residents. International students who are interested in applying to this program must secure an outside grant or fellowship.

Vivien Thomas PhD Scholars

The Vivien Thomas Scholars Initiative (VTSI) is an endowed fellowship program at Johns Hopkins for STEM PhD students. It provides full tuition, stipend, benefits, targeted mentoring, and professional development. Students who have attended a historically black college and university or other minority serving institution for undergraduate study are eligible. Click the following link for more information:  VTSI Flyer

There is no additional application. To be considered for the VTSI, all components of the PhD application, including supplemental components, special VSTI questions, and letters of recommendation must be completed by December 1, 2023, even if the program deadline is later.

Relocation Funds for Incoming Graduate Students

We recognize that it can be financially burdensome to relocate to a new city to attend a Ph.D. program. Students who are accepted to Ph.D. programs at JHU can apply to receive a $1500 need-based grant to offset the costs of relocating to JHU.

These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their Ph.D. program.

This is not a merit-based grant. Applications will be evaluated solely based on financial need.

Click  here  for more information.

Program Requirement

In addition to completing an original research thesis, all Johns Hopkins University PhD candidates are required to pass the Graduate Board oral exam. CBI students are required to meet the following additional requirements:

- Present and defend an original  research proposal  in the CBI Forum during the fall semester of the second year in residence - Complete 3  Research Rotations  in the first year in residence - Complete 8  graduate-level courses

CBI Forum (030.613/614)

CBI students and faculty meet monthly in a forum. The forum hosts:

CBI preceptor presentations – Fall semester First-year student rotation presentations – Throughout the year Third-year Research Proposal Seminars – Fall semester Advanced-year student research updates – Spring semester Seminars presented by distinguished visitors – Throughout the year.  

Third-Year Research Proposal Seminar

As a third-year CBI student, students will prepare and present a research proposal in an area that is unrelated to their dissertation research. The research proposal will help students develop the critical reading, grant proposal writing, and presentation skills that will enhance their success as a research scientist.

Graduate Studies in Chemical Biology

Graduate studies in chemical biology at yale.

The application of chemical concepts and approaches towards a better understanding of biological phenomena and human disease requires a broad multidisciplinary training. Yale University’s Chemical Biology Training Program  draws upon the strengths of several departments to provide interdisciplinary training at the interface of chemistry and biology. Special features of the Chemistry/Biology Interface (CBI) training program include interdepartmental course work and rotation schedule, a Fall Chemical Biology Retreat, a monthly in-house seminar program for discussion of recent results and an annual Chemical Biology Symposium.

The principles of cooperation and collaboration between investigators with diverse expertise and interests are at the heart of Yale’s CBI Graduate Training Program. A friendly, supportive environment, open discussion, shared resources, and multi-lab training are fundamental parts of these studies.

Yale Graduate Studies in Chemical Biology combine the strengths of selected faculty from the  Chemistry Department , the   Molecular, Cellular and Developmental Biology Department , the   Pharmacology Department   , and the   Department of Molecular Biophysics & Biochemistry . Some faculty are part of the   Institute of Biomolecular Design and Discovery . PhD students often join Chemical Biology labs by applying to PhD programs through the   Chemistry Department , or by applying to the   Yale Biological and Biomedical Sciences   program and choosing the  Biochemistry, Quantitative Biology, Biophysics and Structural Biology Track ,   Molecular Cell Biology, Genetics, and Development Track   or the  Molecular Medicine, Pharmacology & Physiology Track .

Chemical Biology

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The Chemical Biology PhD Program is one of the programs in the Harvard Integrated Life Sciences, which facilitates collaboration and cross-disciplinary research. Visit HILS for additional  application instructions .

Chemical Biology is a rapidly growing field that combines the rigor and quantitative aspects of traditional chemistry with the excitement and medical relevance of modern molecular, cellular, organismic, and human biology. You will be encouraged to develop your own ideas, drawing on the expertise available in the community to make exciting new discoveries. You will be able to take full advantage of the unique opportunity to work with a renowned faculty to devise novel approaches to important biological, medical, and chemical problems.

As part of the program, you can interact with the entire campus, including Harvard Medical School, any of the hospital labs, and the Broad Institute. This flexibility enables you to choose the path that most interests you without restriction.

Examples of theses and dissertations completed by students in the program include “All-Optical Neurophysiology in the Photon-1 Regime,” “Biophysical Studies of the LPT Pathway,” and “Chemical Tools to Study Bromodomains in Cancer.”

Graduates of the program have gone into private industry at biotech companies like Novartis, Merck, and Biogen. Others have secured academic positions at prestigious universities like MIT, Stanford University, and Vanderbilt University.

Additional information on the graduate program is available from the Chemical Biology PhD Program  and requirements for the degree are detailed in Policies .

Admissions Requirements

Please review admissions requirements and other information before applying. You can find degree program-specific admissions requirements below and access additional guidance on applying from the Chemical Biology PhD Program .

Academic Background

Typical students have a strong background in one of the disciplines relevant to chemical biology and a strong interest in interdisciplinary research. Be sure to indicate the faculty that interest you in the Faculty section of the application for admission. Students are not required to contact any faculty in advance but are welcome to do so.

Applicants who have been selected for an on-campus interview will be notified directly by the program in Chemical Biology. These visits allow students to meet with faculty and current students to get to know our community and the types of research conducted here. Applicants invited for an interview who reside overseas and cannot visit the Harvard campus may interview remotely.

Standardized Tests

GRE General: Optional

Theses & Dissertations

Theses & Dissertations for Chemical Biology

See list of Chemical Biology faculty

APPLICATION DEADLINE

Questions about the program.

Biology Inspires New Chemical Tools. Chemistry Brings New Understanding to Biological Questions.

The Chemical Biology Graduate Program at UC Berkeley aspires to equip graduate Ph.D. students with the training needed to propel research at the chemistry/biology interface in new and exciting directions while supporting their personal research interests. This interdisciplinary program spans nine departments and over 45 labs .

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News / events, important chemical biology dates for 2020-2021.

Important Dates 1st Rotation: Sept 21 – Nov 23 Poster Session 1: Wed Nov 18  2nd Rotation: Nov 30 – Feb 19 Advisor Check Due: Tuesday Dec 17 (unofficial rotation 3 selection) Poster Session 2: Wed Feb 17 3rd Rotation: Feb 22 – April 30 Poster Session 3: Wed April 28 Thesis Lab Placement: May …

Jennifer Doudna won the 2020 Nobel Prize in Chemistry!

  We are delighted to celebrate Jennifer Doudna winning the Nobel Prize in Chemistry, sharing the award with Emmanuelle Charpentier on CRISPR-Cas9, a transformative method for genome editing! This amazing basic science discovery in the fundamental chemistry of life has opened a whole new world of opportunities to harness this chemical biology technology for the …

Welcome 2020 CBGP Class!

The Chemical Biology Graduate program extends a warm welcome to all incoming CBGP students!  

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Ph.D. Program

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Upcoming Event

• CSB Special Seminar: Steffen Lindert, Ph.D., Wednesday, May 15, 11:00 am!

Advances In Basic Science And Molecular Medicine

The Department of Chemical and Systems Biology explores the frontiers of basic science and molecular medicine, particularly at the crossroads of cellular, chemical, and computational biology. We train Ph.D. students to apply genetic, chemical, cell biological, and quantitative methods to decipher the complex regulatory systems associated with normal physiology and disease states.

Specific research areas include cell signaling pathways, cell cycle control, epigenetics, cell fate specification, and genomic stability. The Chemical and Systems Biology Ph.D. program also emphasizes collaborative learning, and our research community includes scientists trained in molecular biology, cell biology, chemistry, physics, and engineering.

Our Ph.D. program consistently ranks among the top graduate training programs in the world. Most recently the National Research Council named us the top pharmacology-related training program in the United States, based on students’ GRE scores, faculty publications, median time to degree, program requirements, and training resources. The Chemical and Systems Biology graduate program was especially commended for the quality of its research activities.

Why Chemical And Systems Biology?

How do cells achieve directed migration? Why doesn’t a skin cell become a neuron? How do drug-resistant cancers arise and how might they be prevented or overcome? Finding answers to these and other biomedical questions increasingly requires molecular, quantitative, and interdisciplinary approaches.

The Department of Chemical and Systems Biology is uniquely focused on understanding cell biology at the molecular and systems levels, and many of its faculty have expertise in biochemistry, chemistry, physics, and engineering. Developing novel technologies for basic research and translating discoveries into therapeutic strategies are also areas of special interest in the Chemical and Systems Biology community.

Our goal is to train a new generation of scientists with the interdisciplinary skills and creative thinking required to tackle emerging challenges in biomedical research. We invite all interested students to apply to the Chemical and Systems Biology Ph.D. program through the Stanford Biosciences online application form. Applicants whose research interests match well with our scientific mission are encouraged to select Chemical and Systems Biology as their primary home program.

Ph.D. Program

Entering the ph.d. program.

The official course of study in the Ph.D. graduate program begins during the second week of August, one week before the official start of the Fall Semester at Cornell. All incoming Ph.D. students take a series of graduate proficiency exams in Organic, Inorganic, and Physical Chemistry provided by the American Chemical Society (ACS). All Ph.D. students then meet with the Director of Graduate Studies (DGS) and select professors in their area of interest for advice on course selection.

Chemistry and Chemical Biology Ph.D. Program Handbook

Read the Chemistry and Chemical Biology Ph.D. Program Handbook, here .

Ph.D. Coursework

Incoming Ph.D. students generally take three graduate courses during their first semester at Cornell. A minimum grade of B- is required in each course for the student to remain in good standing with the department and the university. An additional three courses are then taken in the spring semester, for a total of six required courses. Depending on a student’s academic background and research interests, one or more of these courses may be taken outside of the Graduate Field of Chemistry & Chemical Biology. Additional courses are often taken by Ph.D. students in the later years of their dissertation work, if they are deemed useful by the student's research advisor and/or special committee (see below). For the full list of courses offered at Cornell, please visit the Class Roster to select the appropriate department and semester.

Finding a Mentor and Laboratory to Conduct Thesis Research

During the first month of the Fall semester, all incoming Ph.D. students are expected to attend a series of research orientation lectures in which the faculty provide an overview of their current research projects. Students are expected to attend research group meetings of faculty of interest, talk to other students and postdoctoral research associates, and discuss potential research projects with at least three faculty members. Students then officially join research groups by November 1.

Special Committee

All Ph.D. students in C&CB are required to choose three or more faculty members to serve as a special committee to represent their major (and minor, if applicable) areas of study. The student’s faculty research advisor serves as chair of the special committee and usually has primary responsibility for directing the graduate student’s research and studies. Degree requirements are kept to a minimum and there are no specific course requirements. The number of formal courses required depends on students' academic background, chosen concentration, and the advice of the special committee.

Every Ph.D. student takes an oral examination for admission to candidacy (A-exam), typically during their second year of graduate study. The A-exam takes place after the student’s coursework has been completed and before the commencement of full-time research. The thesis, which is the final outcome of this research, must constitute an original contribution to chemical knowledge and be defended at a final examination overseen by the special committee (B-exam). The Ph.D. degree is awarded on successful defense of the thesis and students typically take five years to complete the Ph.D. program.

Financial Support

Complete financial support accompanies every offer of admission to the Ph.D. program. Each Ph.D. student is therefore guaranteed at least five years of full financial support as long as he or she makes satisfactory progress toward the Ph.D. degree. This support includes a 12-month stipend, a full tuition award, and health insurance. Financial support comes in the form of teaching assistantships, graduate research assistantships, research fellowships, and several NIH-funded training grant programs, such as the  Chemistry Biology Interface (CBI) Training Program . Eligible applicants are strongly encouraged to seek federally funded fellowships, such as those available from the National Science Foundation (NSF) as well as other government or private agencies.

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Applications for Fall 2024 are now closed.

Applications for Fall 2025 will open in mid August 2024.  All applications and materials will be due no later than December 1, 2024 (11:59PM EST).

The U-M Program in Chemical Biology is one of the largest free-standing interdepartmental programs in the U.S. offering a Ph.D. degree in Chemical Biology. Students receive unparalleled opportunity for research and the flexibility to tailor the program around their own interests.

Participating faculty have appointments in departments including:

• Biological Chemistry
• Medicinal Chemistry
• Pharmacology
• Microbiology and Immunology
• Molecular, Cellular and Developmental Biology
• Molecular and Integrative Physiology
• Computational Medicine
• Bioinformatics 
• Biomedical Engineering

Curriculum & Candidacy

The doctoral program is designed around a flexible curriculum and designed to conclude in five years.

All students take core courses (Chemistry 525; Chemical Biology 510) designed to provide a strong core of the fundamentals of chemical biology. Additional Chemical Biology coursework provides skills in critical analysis and writing. Based on individual interests, the remaining course requirements of the program can be fulfilled from a variety of chemistry, biochemistry or biology classes taught on campus. Hence, a major advantage of the Chemical Biology Program is that it allows students maximum flexibility in their training program. 

Admission to candidacy marks the transition from a largely classroom-based experience to one that is focused on independent research, and occurs before the third year. Post-candidacy regular annual meetings of the student's dissertation committee are held to ensure progress concludes with a dissertation and defense in the fifth year of the program.

The Chemical Biology Program is committed to providing all graduate students with 12 months of financial support in each of the four to five years of their graduate program tenure.  

All students are admitted with full funding (tuition, stipend and benefits) in their first year through Chemical Biology Program fellowships. Additional fellowship opportunities are available for the first year. 

Beyond the first year, students receive aid through a combination of fellowships, research assistantships and teaching assistantships that provide tuition, excellent health and dental benefits and a stipend.  

The program has no teaching requirement, although opportunities are available in the Chemistry Department. 

Also, there are several campus-wide training grants provided by the federal government, which support Chemical Biology graduate students. In most cases, selection for these training programs occurs in the second year of graduate study. The U-M Rackham Graduate School also awards a number of four-year fellowships to students from backgrounds that are historically under-represented in the sciences. 

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TPCB is a leading PhD graduate program in chemical biology, offered jointly by three premier institutions in New York City. We provide an unparalleled combination of world-class faculty, state-of-the-art facilities, and collaborative research opportunities to the next generation of scientific leaders working at the interface of chemistry, biology, and medicine.

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In a recent Nature paper, TPCB student Christian Baca and colleagues in the labs of Prof. Luciano Marraffini and Prof. Dinshaw Patel have discovered a new mechanism of bacterial immunity to phage infection, in which binding of a cyclic adenylate to a novel CRISPR effector called Cam1 arrests host cell functions. [ more ]

TPCB is pleased to partner with the New York Academy of Sciences to present the annual NYAS Chemical Biology Symposium on June 6, 2024. This event is a highlight every year and will feature faculty keynote lectures as well as talks and posters by students and staff scientists from NYC-area labs. [ Register here ]

TPCB students attended the 2023 Career Development Retreat on September 26–27 in the Poconos. The event featured a keynote presentation by Jesse Ausubel, a panel discussion with TPCB alumni, and lots of fun and relaxation. The event was organized by TPCB students Jared Ramsey , Nicolai Tornow , and Nora Zhou . [ more ]

The New York Academy of Sciences will be presenting a symposium on Chemical Biology of Protein Homeostasis: From Fundamentals to Translational Breakthroughs on June 17, 2024. Keynote speakers include Heeseon An (TPCB and MSK), Miklos Bekes, Steve Elledge, Ophir Shalem, and Victoria Kutilek. [ Register here ]

Our Diversity is Our Strength. TPCB has a long-standing commitment to promoting diversity and inclusion at all levels in science. Our outstanding students come from many different backgrounds and the program has zero tolerance for racism or discrimination in any form. We have taken numerous recent efforts to promote these principles. [ more ].

TPCB welcomed the 2023 class of undergraduate summer interns in the Chemical Biology Summer Program (ChBSP) ! These students pursued forefront chemical biology research in TPCB labs, working alongside our students and faculty, as well as participating in career development training. [ more ]

Congratulations to Gianna Stella , who has been recognized for her exceptional contributions to the community with the inaugural TPCB Student Service Award ! Gianna has been a leader in science outreach and mentorship, and a strong advocate for student interests in the graduate schools. [ more ]

The 19th Annual Tri-Institutional Chemical Biology Symposium , sponsored by TPCB, was held on September 6, 2023. Keynote lectures were given by Profs. Danica Fujimori, Viviana Risca, Brian Shoichet, and Jorge Torres. Attendees also enjoyed student talks and a large poster session. Stay tuned for a recap!

TPCB recently welcomed four new faculty to the program, Prof. Arvin Dar at MSK, Prof. Tobias Meyer at Weill Cornell, and Prof. Jiankun Lyu and Prof. Michael Rout at Rockefeller. Their labs bringing exciting new research opportunities in drug discovery and the molecular mechanisms underlying key cellular processes. [ more ]

Alumni Profile : TPCB alumni Dr. Renato Bauer and Dr. Jacqueline Wurst discuss the flexibility of the graduate program, the unique opportunities in the Tri-Institutional research environment, and how their training in chemistry prepared them for their current careers in the pharmaceutical industry at Eli Lilly. [ more ]

Student Profile : TPCB student Chaya Stern describes her non-traditional path to graduate school, where she is now working in Prof. John Chodera’s computational chemistry lab at Sloan Kettering as an NSF Graduate Research Fellow. She also discusses the support she has received from TPCB as a parent. [ more ]

Alumni Profile : TPCB alumna Prof. Niroshika Keppetipola discusses her training experience with mentor Prof. Stewart Shuman and how it prepared her for an independent academic career. Now a faculty member at Cal State Fullerton, her lab studies post-translational regulation of RNA binding proteins in alternative splicing. [ more ]

Faculty Profile : TPCB faculty member Prof. Yael David is a new faculty member at Sloan Kettering. She describe her lab’s approach using protein engineering to study epigenetic regulation and its correlation with disease states. She also discusses her approach to mentoring and her experiences as a woman in science. [ more ]

Alumni Profile : TPCB alumna Dr. Amy Grunbeck Perea developed photoactivatable probes to study G protein-coupled receptors with Prof. Tom Sakmar. After graduating with 9 papers on her CV, she did a postdoc fellowship at Genzyme, then joined Abcam, a biotech company in Cambridge, developing new tools for protein analysis. [ more ]

Faculty Profile : TPCB faculty member Prof. Jue Chen is an HHMI Investigator who studies the structure and function of molecular pumps called ABC transporters that are involved in many important biological processes. She discusses the recent cryo-EM ‘resolution revolution’ and her personalized approach to mentoring her students. [ more ]

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Chemical Biology

Training in chemical biology at MIT enables graduate students to address problems that transcend the traditional boundaries of chemistry and biology. Chemical biologists interrogate molecular processes that occur in human, plant, and microbial cells, as well as in viruses and animals.

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Chemistry and Chemical Biology

Graduate Studies

• Doctor of Philosophy
• Master of Science

The Chemistry and Chemical Biology Graduate Program at UC Davis is world-renowned and represents the full breadth of the discipline, including analytical, biological, inorganic, organic, physical, and theoretical chemistry. The program has over 200 graduate students engaging in research across six areas of study: analytical, chemical biology, inorganic, organic, physical, and chemical physics. We encourage and support graduate research that transcends traditional academic boundaries by bringing together faculty from departments across campus, including the School of Medicine, College of Engineering, College of Biological Sciences and the Genome Center. Students pursuing their Ph.D. receive a competitive salary, tuition and fees paid for and comprehensive health insurance.

Graduate Program Requirements

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Best Global Universities for Biology and Biochemistry in the United States

These are the top universities in the United States for biology and biochemistry, based on their reputation and research in the field. Read the methodology »

To unlock more data and access tools to help you get into your dream school, sign up for the  U.S. News College Compass !

Here are the best global universities for biology and biochemistry in the United States

Harvard university, massachusetts institute of technology (mit), stanford university, university of california berkeley, university of california san francisco, university of california san diego, university of washington seattle, johns hopkins university, university of california los angeles, cornell university.

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• # 1 in Best Universities for Biology and Biochemistry
• # 1 in Best Global Universities

Founded in 1636, Harvard University is the oldest higher education institution in the U.S. The bulk of Harvard's... Read More

• # 2 in Best Universities for Biology and Biochemistry
• # 2 in Best Global Universities

Massachusetts Institute of Technology, founded in 1861, is located in Cambridge, Massachusetts, near Boston. Around... Read More

• # 3 in Best Universities for Biology and Biochemistry
• # 3 in Best Global Universities

Stanford University was founded in 1885 and is located in California’s Bay Area, around 30 miles south of San Francisco... Read More

• # 4 in Best Universities for Biology and Biochemistry
• # 4 in Best Global Universities

The University of California—Berkeley is situated roughly 15 miles from San Francisco in what is known as the Bay Area... Read More

• # 5 in Best Universities for Biology and Biochemistry
• # 16 in Best Global Universities  (tie)

The University of California—San Francisco is a public institution that was founded in 1864. The health sciences-focused... Read More

• # 7 in Best Universities for Biology and Biochemistry
• # 20 in Best Global Universities

The University of California—San Diego is a public institution that was established in 1960. It is located in the La... Read More

• # 8 in Best Universities for Biology and Biochemistry
• # 6 in Best Global Universities

The University of Washington is a public institution that was founded in 1861. The school's oldest and largest campus in... Read More

• # 10 in Best Universities for Biology and Biochemistry
• # 10 in Best Global Universities

Johns Hopkins University is a private institution that was founded in 1876. The school has campuses located in and... Read More

• # 12 in Best Universities for Biology and Biochemistry
• # 14 in Best Global Universities

The University of California—Los Angeles, commonly referred to as UCLA, is a public institution that was founded in... Read More

• # 13 in Best Universities for Biology and Biochemistry
• # 21 in Best Global Universities

Cornell University is a private institution that was founded in 1865. The Ivy League school is located in Ithaca, New... Read More